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Monster Hunter Generations Ultimate features gameplay similar to past titles in the series. The player assumes the role of a hunter who embarks on quests to hunt dangerous creatures. A hunter's abilities are determined by the type of armor and weapons that they wear on a quest, as the hunter otherwise has no intrinsic attributes that affect gameplay. All fourteen weapon types from Monster Hunter 4 Ultimate, ranging from swords, hammers, bows, guns, and lances, are included in Monster Hunter Generations, in addition to the new Prowler mode which allows the player to take the role of a Felyne, a sentient cat-like species. Each weapon has different sets of moves and abilities that can be employed while in the field. Armors grant defensive bonuses to physical and elemental damage, and can boost specific skills and attack types through base attributes and the addition of special decoration gems and talismans.
New in Generations is the ability to transform armor pieces into new gear similar to weapon upgrade paths by upgrading it with materials from broad categories, like bones or ores, and having the ability to upgrade a weapon directly to a more advanced version without the intermediate upgrade steps.